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TAXA . . . . . . . . . . . . . . . . . . . . . . . . .	LUCANIDAE DIPHYLLOSTOMATIDAE PASSALIDAE CHIRONIDAE TROGIDAE GLARESIDAE PLEOCOMIDAE GEOTRUPIDAE BELOHINIDAE OCHODAEIDAE CERATOCANTHIDAE HYBOSORIDAE GLAPHYRIDAE
	SCARABAEIDAE . . . . . . . . . . . .	Aphodiinae Scarabaeinae Pachypodinae Orphinae Allidiostomatinae Dynamopodinae Aclopinae Euchirinae Phaenomeridinae Melolonthinae Rutelinae Dynastinae Cetoniinae