Tribe
GENIATINI Burmeister, 1844:
478
13 genera
330 species and subspecies
BOLAX
Fischer von Waldheim, 1829: 45
Loxopyga Westwood, 1833: Notice 72 (synonym by Ohaus 1918b:
199)
[Gender:
Feminine]
45
species and subspecies
B.
albopilosa Ohaus, 1917: 4
B. albopilosus Ohaus (original spelling by Ohaus)
B. albopilosa Ohaus (justified emendation by Blackwelder
1944: 247)
B. albopilosus Ohaus (unjustified emendation by Frey 1976a:
116)
PERU: Cusco, Ucayali
B.
andicola Burmeister, 1844: 490
B. anticola Erichson, 1847: 100 (unjustified emendation
by Erichson 1847: 100)
PERU: Cusco, Ica
B.
angulata Blanchard, 1851: 235
B. angulatus Blanchard (original spelling by Blanchard)
B. angulata Blanchard (justified emendation by Blackwelder
1944: 247)
B. angulatus Blanchard (unjustified emendation by Frey 1976a:
111)
BRAZIL: Paraná
B.
boliviensis Ohaus, 1898: 63
syn. B. pilosipennis Ohaus, 1917: 5 (synonym by Frey
1976a: 115)
BOLIVIA: La Paz; PERU: Ica, Cusco
B.
buckleyi Ohaus, 1931: 227
ECUADOR
B.
campicola Machatschke, 1974: 247
BRAZIL: Goiás
B.
castaneicollis Burmeister, 1844: 489
BRAZIL: Santa Catarina; COLOMBIA: Bogotá
B.
catharinae Blanchard, 1851: 236
BRAZIL: Santa Catarina
B.
caurana Ohaus, 1931: 228
B. cauranus Ohaus (original spelling by Ohaus)
B. caurana Ohaus (justified emendation by Blackwelder 1944:
247)
B. cauranus Ohaus (unjustified emendation by Frey 1976a:
113)
VENEZUELA
B.
costipennis Ohaus, 1928: 403
Bolax
costipennis Machatschke, 1974: 249. NEW HOMONYM, NEW SYNONYM
BRAZIL: Mato Grosso, Rio de Janeiro, São Paulo
Note:
Machatschke (1974b) is credited with the name B. costipennis (e.g.,
Frey 1976), but Ohaus (1928) properly described the species. Erroneously,
Machatschke (1974b) stated that he was newly describing a species using
Ohaus’ labeled specimens and maintaining Ohaus’ given name
for that species. Machatschke
was unaware that Ohaus (1928) had described the species.
According to these publications, Machatschke and Ohaus used the
same specimens for the type series. According to Article 60.2 (ICZN 1999),
if the rejected junior homonym has one or more potentially valid synonyms,
the oldest of these names is the valid name of the taxon (B. costipennis
Ohaus in this case).
B.
cupreoviridis Ohaus, 1931: 229
PERU: San Martin
B.
disgamia Ohaus, 1917: 5
B. disgamius Ohaus (original spelling by Ohaus)
B. disgamia Ohaus (justified emendation by Blackwelder 1944:
247)
B. disgamius Ohaus (unjustified emendation by Frey 1976a:
113)
PERU: Ica
B.
flaveola Bates, 1888: 295
B. flaveolus Bates (original spelling by Bates)
B. flaveola Bates, 1888: 295 (justified emendation by Blackwelder
1944: 247)
B. flaveolus Bates (unjustified emendation by Frey 1976a:
112)
COSTA RICA; NICARAGUA
B.
flavolineata (Mannerheim, 1829: 60)
Geniates flavolineatus Mannerheim
Leucothyreus flavolineatus (Mannerheim) (new comb. by Laporte
1840: 139)
B.
flavolineatus (Mannerheim) (new comb. with incorrect species-group
gender by Burmeister 1844: 487)
B.
flavolineata (Mannerheim) (justified emendation by Blackwelder 1944:
247)
Loxopyga
bicolor Westwood 1833: Notice 72 (synonym by Burmeister 1844: 487
BRAZIL: Minas Gerais, Rio de Janeiro, São Paulo
B.
foveolata Blanchard, 1851: 236
Bolax foveolatus Blanchard (original spelling by Blanchard)
B. foveolata Blanchard (justified emendation by Blackwelder
1944: 247)
B. foveolatus Blanchard (unjustified emendation by Frey
1976a: 115)
BRAZIL: Mato Grosso; FRENCH GUIANA
B.
gaudichaudi Blanchard, 1851: 236
BRAZIL
B.
glabripennis Ohaus, 1917: 7
PERU: Loreto, Huanuco, La Libertad, San Martin
B.
goyana Ohaus, 1917: 8
B. goyanus Ohaus (original spelling by Ohaus)
B. goyana Ohaus (justified emendation by Blackwelder 1944:
248)
B. goyanus Ohaus (unjustified emendation by Frey 1976a:
114)
BRAZIL: Goiás
B.
hirtula Burmeister, 1844: 491
B. hirtulus Burmeister (original spelling by Burmeister)
B. hirtula Burmeister (justified emendation by Blackwelder
1944: 248)
B. hirtulus Burmeister (unjustified emendation by Frey 1976a:
116)
COLOMBIA: Bogotá
B.
incogitata Dohrn, 1883: 427
B. incogitatus Dohrn (original spelling by Dohrn)
B. incogitata Dohrn (justified emendation by Blackwelder
1944: 248)
B. incogitatus Dohrn (unjustified emendation by Frey 1976a:
109)
PERU: Arequipa
B.
magna Bates, 1888: 295
B. magnus Bates (original spelling by Bates)
B. magna Bates (justified emendation by Blackwelder 1944:
248)
B. magnus Bates (unjustified emendation by Frey 1976a: 110)
syn.
Bolax vittatus Casey, 1915: 105 (synonym by Ratcliffe 2002: 26) invalid synonym
PANAMA:
Bocas del Toro, Canal Zone, Chiriqui, Darien; COSTA RICA
Note:
Ratcliffe (2002) synonymized Bolax vittatus Casey in error. Ratcliffe studied Casey’s type specimen and determined
that it was congeneric with Trizogeniates foveicollis Ohaus. However, in a “senior moment” (pers. comm. to MLJ,
Oct. 2004), Casey’s name and the synonymy were misplaced in Ratcliffe’s
(2002) publication. Villatoro
(2002) correctly synonymized B. vittatus Casey with T. foveicollis.
B.
matogrossensis Ohaus, 1917: 3
BRAZIL: Mato Grosso
B.
mutabilis Burmeister, 1844: 489
syn. B. nitens Blanchard, 1851: 236 (synonym by Ohaus
1917: 9)
BRAZIL: Espirito Santo, Rio de Janeiro, São Paulo
B.
nigriceps Ohaus, 1917: 6
BOLIVIA; PERU: Ica
B.
oberthuri Ohaus, 1903: 235
ECUADOR: Loja, Tungurahua
B.
palliata palliata Burmeister, 1844: 491
B. palliatus Burmeister (original spelling by Burmeister)
B. palliata Burmeister (justified emendation by Blackwelder
1944: 248)
B. palliatus Burmeister (unjustified emendation by Frey
1976a: 110)
VENEZUELA: Mérida
B.
palliata fuscipennis Ohaus, 1917: 4
B. palliatus infuscipennis Ohaus (original spelling by Ohaus)
B. palliata infuscipennis Ohaus (justified emendation by
Blackwelder 1944: 248)
B. palliatus fuscipennis Ohaus (unjustified emendation
by Frey 1976a: 110)
VENEZUELA: Zulia
B.
palliata infuscata Ohaus, 1917: 4
B. palliatus infuscatus Ohaus (original spelling by Ohaus)
B. palliata infuscata Ohaus (justified emendation by Blackwelder
1944: 248)
B. palliatus infuscatus Ohaus (unjustified emendation by
Frey 1976a: 110)
COLOMBIA:
Atlantico
B.
palliata unicolor Benderitter, 1925: 248
COLOMBIA
B.
phalerata Burmeister, 1844: 488
B. phaleratus Burmeister (original spelling by Burmeister)
B. phalerata Burmeister, 1844: 488 (justified emendation
by Blackwelder 1944: 248)
B. phaleratus Burmeister (unjustified emendation by Frey
1976a: 112)
syn. B. tibialis Blanchard, 1851: 235 (synonym by
Frey 1976a: 112)
syn. B. salticola Ohaus, 1908: 262 (synonym by Frey
1976a: 112)
BRAZIL: Rio de Janeiro, São Paulo
B.
pulla (Latreille, 1833: 13)
Melolontha pullus Latreille (original spelling by Latreille)
B.
pullus (Latreille) (new comb. with incorrect species group gender
by Burmeister 1844: 491)
B. pulla (Latreille) (justified emendation by Blackwelder
1944: 248)
ECUADOR: Pichincha
B.
rutila Erichson, 1847: 100
B. rutilus Erichson (original spelling by Erichson)
B.
rutila Erichson (justified emendation by Blackwelder 1944: 248)
B.
rutilus Erichson (unjustified emendation by Ohaus 1952: 10; Frey 1976a:
113)
PERU: Ica
B.
saucia Ohaus, 1917: 9
B. saucius Ohaus (original spelling Ohaus)
B. saucia Ohaus (justified emendation by Blackwelder 1944:
248)
B. saucius Ohaus (unjustified emendation by Frey 1976a:
114)
BRAZIL: Minas Gerais
B.
sculpticollis Frey, 1976: 117
BRAZIL: Espirito Santo, Minas Gerais
B.
squamulifera Blanchard, 1851: 236
B. squamuliferus Blanchard (original spelling by Blanchard)
B. squamulifera Blanchard (justified emendation by Blackwelder
1944: 248)
B. squamuliferus Blanchard (unjustified emendation by Frey
1976a: 116)
FRENCH GUIANA
B.
sulcicollis (Germar, 1824: 124)
Melolontha sulcicollis Germar
B. sulcicollis (Germar) (new comb. by Burmeister 1844: 489)
BRAZIL: Bahia, Espirito Santo
B.
sulcipennis Ohaus, 1928: 404
BRAZIL: Bahia
B.
tacoaraphaga Ohaus, 1908: 257
BRAZIL: Espirito Santo, Minas Gerais, Rio de Janeiro
B.
variolosa Ohaus, 1917: 4
B. variolosus Ohaus (original spelling by Ohaus)
B. variolosa Ohaus (justified emendation by Blackwelder
1944: 248)
B. variolosus Ohaus (unjustified emendation by Frey 1976a:
110)
COLOMBIA: Bogotá, Valle del Cauca; PANAMA: Veraguas
B.
vauriae Frey, 1976: 110
BRAZIL
B.
vittipennis (Laporte, 1840: 139)
Aulacodus vittipennis Laporte
B. vittipennis (Laporte) (new comb. by Blanchard 1851: 236)
syn. B. leucogaster Burmeister, 1844: 492 (synonym
by Burmeister 1855: 534)
BRAZIL
and FRENCH GUIANA (Blackwelder 1944); FRENCH GUIANA and GUYANA (Machatschke
1972-1974); BRAZIL: Mato Grosso, São Paulo (Frey 1976).
*B.
zoubkovii Fischer von Waldheim, 1829: 45
B.
zubkoffi Fischer von Waldheim (unjustified emendation by Ohaus 1908a:
254-256, 262)
B. zoubkoffi Fischer von Waldheim (unjustified emendation by
Ohaus 1917: 8)
syn.
B. westwoodi Laporte, 1840: 140 (original spelling by Laporte)
B. westwoodii
Laporte (synonym and unjustified emendation by Burmeister 1844: 488)
BRAZIL: Minas Gerais, São Paulo
EUNANUS Ohaus, 1909: 442
[Gender:
Masculine]
2
species
*E.
murinus Ohaus, 1909: 442
ARGENTINA: Buenos Aires, Santa Fé
E.
podtiaguini Gutiérrez, 1951: 121
ARGENTINA: Chaco
EVANOS
Laporte, 1840: 140
[Gender:
Masculine]
1
species
*E.
villatus Laporte, 1840: 140
BRAZIL: Distrito Federal, Espirito Santo, Rio de Janeiro
GENIATES Kirby, 1819: 401
[Gender:
Masculine]
39
species
G.
affinis Camerano, 1878: 237
BRAZIL
G.
balzapambae Ohaus, 1917: 46
ECUADOR: Bolivar; PERU
*G.
barbatus Kirby, 1819: 403
BRAZIL
G.
bituberculatus Camerano, 1878: 235
G. bituberculata Camerano (unjustified emendation by Blackwelder
1944: 249)
BRAZIL
G.
borellii Camerano, 1894: 1
G. bornelli Camerano (lapsus by Machatschke 1972: 133)
ARGENTINA: Tucuman
G.
castaneus Burmeister, 1844: 511
G. castanea Burmeister (unjustified emendation by Blackwelder
1944: 249)
BRAZIL: Rio de Janeiro
G.
clavipalpus Burmeister, 1844: 512
BRAZIL
G.
convexus Ohaus, 1917: 47
G. convexa Ohaus (unjustified emendation by Blackwelder
1944: 249)
BRAZIL
G.
corniger Ohaus, 1931: 252
BRAZIL: São Paulo
G.
cornutus Burmeister, 1844: 513
G. cornuta Burmeister (unjustified emendation by Blackwelder
1944: 249)
BRAZIL: Rio de Janeiro
G.
cylindricus Burmeister, 1844: 514
G. cylindrica Burmeister (unjustified emendation by Blackwelder
1944: 250)
ARGENTINA; BRAZIL: Rio Grande do Sul
G.
distans Burmeister, 1844: 508
BRAZIL
G.
ferrugatus Mannerheim, 1829: 62
BRAZIL:
Minas Gerais
G.
flaviventris Frey, 1976b: 353
PANAMA: Panama
G.
fuscescens Camerano, 1878: 237
T.
fuscescens (Camerano) (new comb. by Ohaus 1917: 39)
G.
fuscescens Camerano (new comb. by Ohaus 1918b: 206)
BRAZIL
G.
fuscicollis Ohaus, 1917: 50
BRAZIL
G.
immaculatus Camerano, 1878: 236
L. immaculatus (Camerano) (new comb. by Ohaus 1917: 28)
L. immaculata (Camerano) (unjustified emendation by Blackwelder
1944: 249)
G. immaculatus Camerano (new comb. by Martinez 1964a: 433)
BRAZIL
G.
impressicollis Ohaus, 1911: 559
ARGENTINA;
BRAZIL; PARAGUAY: San Pedro, Central
G.
impressifrons Lucas, 1857: 135
BRAZIL: Minas Gerais
G.
incertus Camerano, 1878: 237
G. incerta Camerano (unjustified emendation by Blackwelder
1944: 249)
BRAZIL
G.
inconstans Ohaus, 1931: 253
BRAZIL: Espirito Santo
G.
isthmicus Ohaus, 1931: 254
G. istmicus Ohaus (lapsus by Ohaus 1931: 255)
G. isthmica Ohaus (unjustified emendation by Blackwelder
1944: 249)
PANAMA
G.
leptopus Frey, 1976: 351
BOLIVIA; BRAZIL: Mato Grosso; CHILE: Santiago
G.
marginicollis Ohaus, 1917: 48
T. marginicollis (Ohaus) (new comb. by Martínez 1956:
8)
G. marginicollis Ohaus (new comb. by Martínez 1965:
9)
ARGENTINA: Entre Rios; BRAZIL: Alagoas, Rio Grande do
Sul, Santa Catarina
G.
multicornis Camerano, 1878: 236
BRAZIL
G.
octavius Ohaus, 1924: 185
G. octavia Ohaus (unjustified emendation by Blackwelder
1944: 250)
BRAZIL
G.
palliatus Ohaus, 1917: 46
G. palliata Ohaus (unjustified emendation by Blackwelder
1944: 250)
ECUADOR: Esmeraldas
G.
pallidus Burmeister, 1844: 512
G. pallida Burmeister (unjustified emendation by Blackwelder
1944: 250)
BRAZIL: Rio de Janeiro
G.
punctipennis Ohaus, 1917: 47
BRAZIL
G.
relictus Ohaus, 1931: 257
G. relicta Ohaus (unjustified emendation by Blackwelder
1944: 250)
BRAZIL: São Paulo
G.
rufescens Lucas, 1857: 136
BRAZIL: Goiás
G.
rugicollis Lucas, 1857: 135
G. ruficollis Lucas (lapsus by Machatschke 1972: 356)
BRAZIL
G.
rugosus Camerano, 1878: 236
G. rugosa Camerano (unjustified emendation by Blackwelder
1944: 250)
BRAZIL
G.
sericeus Burmeister, 1855: 535
G. sericea Burmeister (unjustified emendation by Blackwelder
1944: 250)
BRAZIL: Rio de Janeiro
G.
spurius (Curtis, 1845: 450)
Leucothyreus spurius Curtis
G. spurius (Curtis) (new comb. by Burmeister 1855: 535)
G. spuria (Curtis) (unjustified emendation by Blackwelder
1944: 250)
BRAZIL
G.
subsericeus Ohaus, 1917: 48
G. subscriceus Ohaus (print error by Ohaus 1918b: 206)
G. subsericea Ohaus (unjustified emendation by Blackwelder
1944: 250)
BRAZIL: Minas Gerais
G.
truquii Camerano, 1878: 236
BRAZIL
G.
verticalis Burmeister, 1844: 512
BRAZIL
G.
werneri Ohaus, 1931: 255
COLOMBIA
GENIATOSOMA
Costa Lima, 1940: 62
[Gender:
Neuter]
3
species
G.
matilei Lacroix, 2000: 201
BRAZIL: Amazonas
G.
nigrum (Ohaus, 1917: 49)
Geniates niger Ohaus
Geniatosoma
niger (Ohaus) (new comb. with incorrect species group gender by Costa
Lima 1940: 62)
Geniatosoma nigra (Ohaus) (unjustified emendation by Martínez
1977: 12)
Geniatosoma nigrum (Ohaus) (justified emendation by Lacroix
2000: 200)
FRENCH GUIANA
G.
lindemannae (Frey, 1976: 354)
Geniates lindemannae Frey. NEW COMBINATION
syn. Geniatosoma fernandezi Martínez, 1977:
15. NEW SYNONYM
BRAZIL: Amazonas; VENEZUELA: Amazonas
Note:
Based on our examination of type specimens of Geniates lindamannae
Frey and Geniatosoma fernandezi Martínez, these species
are conspecific. We transfer G. lindamannae to Geniatosoma
and synonymize G. fernandezi with G. lindamannae.
HETEROGENIATES Ohaus, 1909: 444
[Gender:
Masculine]
1
species
*H.
bonariensis Ohaus, 1909: 444
ARGENTINA:
Buenos Aires, Tucuman
LEUCOTHYREUS Macleay, 1819: 145
syn. Aulacodus Eschscholtz, 1822: 22 (synonym by
Burmeister 1844: 493)
syn. Bolaxoides Laporte, 1840: 140 (synonym by Ohaus
1918b: 201)
[Gender:
Masculine]
164
species and subspecies
L.
acanthurus Ohaus, 1917: 20
BRAZIL:
Espirito Santo
L.
acuminatus Ohaus, 1917: 24
BRAZIL: São Paulo
L.
aeneiceps Kirsch, 1871: 370
COLOMBIA: Bogotá
L.
albertus Ohaus, 1918: 352
COLOMBIA: Bogotá
L.
albopilosus Ohaus, 1917: 10
BRAZIL: Espirito Santo, Acre, Mato Grosso, Rio de Janeiro,
São Paulo
L.
aloysius Ohaus, 1918: 350
FRENCH GUIANA
L.
alvarengai Frey, 1976: 349
BRAZIL: Mato Grosso
L.
amandus Ohaus, 1918: 354
FRENCH GUIANA
L.
ambrosius Ohaus, 1918: 355
BRAZIL: Mato Grosso
L.
anachoretus Erichson, 1848: 563
L. anachoreta Erichson (original spelling by Erichson)
GUYANA
L.
anaemicus Ohaus, 1918: 359
ECUADOR
L.
baeri Ohaus, 1917: 25
PERU: Tumbes
L.
bakeri Casey, 1915: 106
BRAZIL: Para
L.
bohlsi Ohaus, 1918: 351
PARAGUAY: Central
L.
bucki Machatschke, 1974: 143
BRAZIL:
Rio Grande do Sul
L.
calcaratus Ohaus, 1917: 24
BRAZIL: Rio Grande do Sul
L.
campestris Burmeister, 1855: 534
BRAZIL: Minas Gerais
L.
camposseabrai Martínez, 1962: 119
BRAZIL: Amazonas
L.
cavifrons Burmeister, 1844: 503
BRAZIL
L.
cayapo Ohaus, 1931: 229
BRAZIL: Goiás
L.
cephalotes Casey, 1915: 105
COSTA RICA
L.
chalceus Blanchard, 1851: 238
BRAZIL: Minas Gerais, São Paulo
L.
chiriguanus Ohaus, 1917: 18
BOLIVIA: La Paz
L.
chloroticus Ohaus, 1918: 359
BRAZIL
L.
clemens Ohaus, 1918: 363
BRAZIL: Amazonas
L.
clodius Ohaus, 1931: 230
BRAZIL: São Paulo
L.
cominius Ohaus, 1931: 232
BRAZIL
L.
conquisitor Ohaus, 1924: 184
BOLIVIA: La Paz
L.
costatus costatus Burmeister, 1844: 498
syn. L. rugipennis Blanchard, 1851: 238 (synonym
by Ohaus 1931: 233)
ARGENTINA; BRAZIL; URUGUAY
L.
costatus chaconus Ohaus, 1931: 233
ARGENTINA: Chaco
L.
cribratipennis Blanchard, 1851: 238
ARGENTINA; BRAZIL: Mato Grosso
L.
crispinus Ohaus, 1918: 353
FRENCH GUIANA; SURINAM
L.
cruralis Ohaus, 1931: 234
BRAZIL: Amazonas
L.
cupripennis Ohaus, 1918: 357
ECUADOR: Pastaza
L.
cuyabanus Ohaus, 1917: 27
BRAZIL: Mato Grosso
L.
cyanescens Blanchard, 1851: 238
BOLIVIA:
Santa Cruz
L.
damasus Ohaus, 1918: 356
VENEZUELA: Distrito Federal
L.
davisi Ohaus, 1908: 407
ECUADOR: Chimborazo
L.
decolor Ohaus, 1917: 14
COLOMBIA; VENEZUELA
L.
demetrius Ohaus, 1918: 358
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